The commercial, titled "Battle Up" follows the evolution of a female Marine.
She is seen as a schoolgirl standing up to bullies, on a rugby pitch, under fire in combat, and as a veteran helping the homeless.
The service said it celebrates the Marines' "fighting spirit".
In the advert, the protagonist tackles an obstacle course through icy water as the narrator intones: "No one knows where it comes from. Why some have it, and some don't. It's the fighting spirit, and it needs to be fed.
