The prop had not been expected to be part of Eddie Jones' squad for the Twickenham encounter, having missed England's win over Ireland after travelling to Tonga to visit a seriously ill relative.
However, he is included in Jones' 34-man party as England begin preparations for the visit of Wayne Pivac's team.
Exeter Chiefs scrum-half Jack Maunder has been selected, giving Jones a third option at number nine behind Ben Youngs and Will Heinz, while wing Anthony Watson and hooker Luke Cowan-Dickie were also named in the squad.
England bounced back from their opening-round defeat to France with victories over Scotland and Ireland.
They trail leaders France - who face Scotland at Murrayfield on Sunday - by four points and know defeat to Wales could spell the end of their hopes of regaining the Six Nations title.
Photo Getty Images Caption: England prop Mako Vunipola